When a gene colour rule states it has to be natural, it may mean a couple of different things!
It can be any of the following: Any natiural tones, ane colour from the base coat swatch (excluding malenism and leucism unless those genes are prsent), or a longter/darker shade of the coat itself. The hue, however, is not allowed to change.
Below, you can see what we consider natural tones, as well as a cisual on what you may change when it comes to different shades of the base coat.